We know that they howl quite often after meals or when a team leaves the yard. They howl when they hear a distant neighbor howl. Of course, they howl when they are "coaxed" to howl by Aliy or Allen.
All in all, they seem to love howling. Their tails are naturally wagging and their heads are thrown back with ecstasy. Some dogs stay in their houses, some jump to attention. Regardless, of what it all means .... it's a lovely sight and sound ..... for us mere humans.